A seed contains the
embryo from which a new plant will grow under proper
conditions. Seeds also usually contain a supply of stored
food and is wrapped in the seed coat or testa. Seeds
are very diverse in size. The dust-like orchid seeds
are the smallest with about one million seeds per gram.
Orchid seeds have immature embryos and no significant
food reserves. They are myco-heterotrophs, depending
on mycorrhizal fungi for nutrition during germination.
At over 20 kg, the largest seed is the coco de mer.
The embryo has one cotyledon or seed leaf in monocotyledons,
two cotyledons in almost all dicotyledons and two
or more in gymnosperms. The radicle is the embryonic
root. The plumule is the embryonic shoot. The embryonic
stem above the point of attachment of the cotyledon(s)
is the epicotyl. The embryonic stem below the point
of attachment is the hypocotyl.
In angiosperms, the stored food begins as a tissue
called the endosperm, which is derived from the parent
plant via double fertilization. The usually triploid
endosperm is rich in oil or starch and protein. In
gymnosperms, such as conifers, the food storage tissue
is part of the female gametophyte, a haploid tissue.
In some species, the embryo is embedded in the endosperm
or female gametophyte, which the seedling will use
upon germination. In others, the endosperm is absorbed
by the embryo as the latter grows within the developing
seed, and the cotyledons of the embryo become filled
with this stored food. At maturity, seeds of these
species have no endosperm and are termed exalbuminous
seeds. Some exalbuminous seeds are bean, pea, oak,
walnut, squash, sunflower, and radish. Seeds with
an endosperm at maturity are termed albuminous seeds.
Most monocots (e.g. grasses and palms) and many dicots
(e.g. brazil nut and castor bean) have albuminous
seeds. All gymnosperm seeds are albuminous.
The seed coat develops from the tissue, the integument,
originally surrounding the ovule. The seed coat in
the mature seed can be a paper-thin layer (e.g. peanut)
or something more substantial. The seed coat helps
protect the embryo from mechanical injury and from
drying out.
The seeds of angiosperms are contained in a hard
or fleshy structure called a fruit. Gymnosperm seeds
begin their development "naked" on the bracts
of cones, although the seeds do become covered by
the cone scales as they develop. An example of a hard
fruit layer surrounding the actual seed is that of
the so-called stone fruits.
Some seeds have an appendage on the seed coat such
an aril or an elaiosome or hairs. The hilum is the
scar on the seed coat where the seed was attached
to the ovary wall by the funiculus.
In order for the seed coat to split, the embryo must
imbibe, which causes it to swell, splitting the seed
coat. However, the nature of the seed coat determines
how rapidly water can penetrate and subsequently initiate
germination. For seeds with a very thick coat, scarification
of the seed coat may be necessary before water can
reach the embryo. Examples of scarification include:
gnawing by animals, freezing and thawing, battering
on rocks in a stream bed, or passing through an animal's
digestive tract. In the latter case, the seed coat
protects the seed from digestion, while perhaps weakening
the seed coat such that the embryo is ready to sprout
when it gets deposited far from the parent plant.
In species with thin seed coats, light may be able
to penetrate into the dormant embryo. The presence
of light or the absence of light may trigger the germination
process, inhibiting germination in some seeds buried
too deeply or in others not buried in the soil. Abscisic
acid is usually the growth inhibitor in seeds.